Oenocytes, a special cell type, often are present between the basal region of the epidermal cells and the basal lamina, or they may adhere to the hemolymphal surface of the basal lamina. Electron microscope studies show that they have a highly developed smooth endoplasmic reticulum, characteristic of cells engaged in hydrocarbon synthesis. Oenocytes synthesize hydrocarbon waxes, which are transferred to the wax layer of the epicuticle, presumably via the epidermal cells and the procuticular pore canals. It has been suggested that the oenocytes also provide lipids to the outer and inner epicuticle as well as to the sclerotized regions of the exocuticle.

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